Back to other Lectures

Unit 6. Ecological Systems

Lecture 27. Populations and Communities

1.  Objectives

1.1.  To present factors that determine population density, distribution and change

1.2.  To describe how communities are organized and how they develop and diversify

2.   Introduction to Ecology

2.1.  Lectures

bulletPopulations and Communities
bulletEcosystems
bulletBiosphere
bulletHuman Impact on Biosphere

2.2.  Some Definitions

2.2.1.  Ecology - interaction of organisms and their environments

2.2.2.  Population - several members of the same species within a defined geographic area

2.2.3.  Community - several species that interact within a defined area

2.2.4.  Ecosystem (27-1) - a system involving interactions of several communities

2.2.5.  Biosphere - global ecosystem

 

3.  Populations

3.1.  Are Units of Structure and Function

3.2.  Population Size and Distribution are Important to Ecosystems

3.2.1.  Often the concentration is more important than numbers

3.2.2.  Individuals are distributed uniformly, randomly, or in clumps (27-2)
bulletexamples (27-3) (tussock grass, crocus, and flamingos)

3.2.3.  Clumping or aggregation is most common
bulletenvironmental conditions are seldom uniform
bulletreproductive patters favor clumping (e.g. vegetative reproduction)
bulletbehavioral patterns often lead to groups (27-4)   (e.g. gannet colony clumping favors protection and reproduction)

3.3.  Populations Under Ideal Conditions Show Exponential Growth (27-5)

3.3.1.  Equation

I = rN
bulletI = population growth rate
bulletr = intrinsic rate of increase = (births - deaths) per individual
bulletN = population size (number of individuals)

3.3.2.  A housefly beginning to breed in April would increase to 2 X 1020 by August in an unlimited environment (would cover 10 cm of earth's land surface)

3.3.3.  In the U.S. the birth rate is 16 per 1000 and death rate is 8.5 per 1000 so:
bulletr = 0.016 - 0.0085 = 0.0075
bulletgrowth rate is 0.75% per year or the population will double in 93 years

3.4.  Most Populations Exhibit an S-Shaped Growth Curve (27-6)

3.4.1.  Carrying capacity (K) - maximum number of population over a sustained period

3.4.2.  Growth of sheep in South Australia (27-7)

3.5.  Some Populations Exhibit Unstable Growth Curves

3.5.1.  boom and bust (27-8) (e.g. pea aphids in an alfalfa field, influenced by weather and season)

3.5.2.  wild fluctuation independent of environmental conditions (27-9) (e.g., rmax >2, population overshoots k approaches 3 and is erratic)

3.5.  Mortality and Survivorship Influence Population Growth

3.5.1.  Plot showing three types (27-10)
bulletlow infant mortality (humans)
bulletmoderate infant mortality (hydra)
bulletinfant mortality high (oysters)

3.6.  Age Distribution Influences Population Growth

3.6.1.  Sweden (27-11) - birth rate = death rate; population is not growing

3.6.2.  U.S. (27-12) - size of youngest group is equal to reproductive group; population will continue to increase until year 2000; note the baby boom bulge

3.6.3.  India (27-13) -  high birth rate; population will increase for next two generations

3.6.4.  Growth of world population (27-14)

3.7.  How Populations are Regulated

3.7.1.  Predation, disease, and parasitism
bulletdensity dependent
bulletexample of predator prey (27-14)
bulletmoose/wolves (27-15)

3.7.2.  Competition
bulletintraspecific - same species - e.g. flowers planted too close
bulletinterspecific - different species -
bulletprinciple of limiting similarity - no two species can occupy the same niche at the same time
bulletniche - ecological role of a species in a community

example - niches of three closely related warblers (27-16)

3.7.3.  Behavioral and Physiological Changes
bulletcooperative hunting of dogs (27-17)
bulletlocusts migrate in phase (food depletion leads to swarming to find food) (27-18)
bulletsocial behavior of ants and bees (27-19)

3.7.4.  Concept of limiting factors - only one factor can be limiting at time

 

4. Communities

4.1.  Succession

4.1.1.  Primary succession - communities established in newly formed habitats, e.g. sand dunes; bare rock

4.1.1.1.  successions in ponds (Presque Isle, PA, a penisula in Lake Erie)
bulletnewly formed pond (27-20)
bullet2-years low vegetation, cottonwood saplings (27-21)
bullet50-years, choked with weeds, cottonwoods (27-22)
bullet150-200 years, meadow (27-23)

4.1.1.2.  succesional stages at southern Lake Michigan (27-24)

4.1.2.  Secondary succession - communities are reestablished where another community once existed
bullete.g. abandoned railway (27-25)

4.1.3.  Bird Succession on abandoned farmland in Georgia (27-26)

4.1.4.  Succession is characterized by a number of trends
bulletfirst species are rapid growers capable of surviving a harsh environment; later species are slower growing, more competitive
bulletspecies composition changes regularly and more rapidly in early stages
bulletnumber of species increases rapidly at first and then stabilizes (27-27)
bulletsuccessional species may alter environment and make it more favorable for competitors
bulletgross primary productivity increases until it reaches a stable high (27-28)
bulletstore of inorganic nutrients in soil and organism of ecosystem increases and a larger proportion is held in tissues of plants
bullettotal biomass and non-living organic matter increase
bulletsize of plants increase
bulletcommunities become more diverse and complex.

4.1.5. Climax community (27-29) - when energy flow and biomass reach equilibrium, i.e. total respiration equals gross primary productivity, e.g. /sequoia forest (some trees are 2,000 years old)

 

8.  Critical Concepts or Questions

8.1.  Define population size, density, and distribution andage structure.

8.2.  Why do populations not restricted in some way grow exponentially?

8.3.  Define carrying capaacity and describe its effect as evidenced by a logistic growth pattern.

8.4.  What are some limiting factors for growth of microbes, plants, and animals?

8.5.  At the present growth rate, how long wil it take before the human population has another billion individuals added to it? 

8.6.  How did earlier human populations expand steadily into new environments?  How did they increase the carrying capacity of their habitats?  How have they avoided some of the limiting factors on population growth?    Are their methods sustainable?

8.7.  What is the difference between the habitat and nich of a species?

8.8.  Why is it difficult to define "the human habitat?"

8.9.  Describe competitive exclusion.  How might two species that compete for the same resource coexist?

8.10.  Define the difference between predator and parasite.

8.11.  Define primary and secondary succession.  Give specific examples of each.

8.12.  What is a climax community?  How does the climax-pattern model help explain its structure?

9.  Vocabulary

biotic potential carrying capacity community
demographic transition model ecology life table
density-dependent controls ecosystem population
density-independent controls habitat limiting factor
exponential growth J-shaped curve logistic growth
zero population growth survivorship curve S-shaped curve

 

camouflage climax community coevolution
commensalism community habitat
competitive exclusion interspecific competition predator
mimicry mutualism niche
parasite parasitism predation
pioneer species secondary succession prey
primary succession resource partitioning symbiosis

10.  Webpage Links

Microbial Ecology - Digital Learning Center for Microbial Ecology, Michigan State University ** Microbe Zoo - images and habitats **, plus Microbes in the News - stories, Meet the Scientists - profiles, Microbial Ecology Resources - listings.


11.  Illustration Documentation